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Anne McElvoy talks to the biologist Ken Thompson who dismisses attempts to control invasive species and questions the veracity of dividing plants and animals into 'native' and 'alien'. However the Director of the Kew Innovation Unit Monique Simmonds warns that alien pests and diseases can have a devastating effect on much-loved plants, and that it's vital to maintain and support diverse environments. The farmer John Lewis-Stempel records the passage of the seasons in his account of the life of an English meadow and he laments the decline of some of his favourite birds from his childhood. The woolly mammoth used to be native in Europe before it became extinct, and the palaeontologist Victoria Herridge confounds expectations by identifying the smallest mammoth ever known to have lived. Producer: Katy Hickman.

Tom Sutcliffe asks whether it's better to lead from the front, or advise from the side-line. The Deputy Chairman of Saatchi & Saatchi, Richard Hytner celebrates the latter: those who wield influence and authority away from the limelight. Heather Rabbatts has experience of being a Deputy and a Chief Executive in both politics and business. The academic Archie Brown looks back at the history of political leadership and questions whether strong leaders are the most successful and admirable, while Tony Blair's former chief of staff, Jonathan Powell turns to Machiavelli's The Prince for a primer on the art of government. Producer: Katy Hickman.

Anne McElvoy talks to the poet Simon Armitage about his dramatisation of The Last Days of Troy. His play, based on Homer's epic, reveals how cycles of conflict and revenge, pride and self-deception continue throughout history. Greek myth is at the heart of a new opera, Thebans, in which the playwright and poet Frank McGuinness draws on the tragedy of the mythical monarch Oedipus and his daughter Antigone. Natalie Haynes explores what happens when troubled teenagers become enthralled by Greek tales of cruel fate and bloody revenge in her debut novel, while Kenan Malik goes on a quest for a moral compass. Producer: Katy Hickman.